王文采 杨宗宗



摘 要:该文描述了在新疆维吾尔族自治区发现的毛茛科翠雀花属三新种,包括隶属于密花翠雀花组高翠雀花亚组的二新种,即布尔津翠雀花(Delphinium buerjinense W. T. Wang & Z. Z. Yang)和尼勒克翠雀花(D. nilekeense W. T. Wang & Z. Z. Yang),以及隶属于翠雀花组须花翠雀花亚组的一新种,即哈巴河翠雀花(D. habaheense W. T. Wang & Z. Z. Yang);此外,还分别描述了此三新种与近缘种的区别特征。
关键词: 毛茛科, 翠雀花属, 新种, 新疆
中图分类号:Q949
文献标识码:A
文章编号:1000-3142(2021)03-0327-07
收稿日期:2021-01-03
基金项目:中国科学院植物研究所资助王文采院士科研活动经费(110100PZ07) [Supported by the Fund for WANG Wen-tsai Academician Scientific Research of Institute of Botany, Chinese Academy of Sciences (110100PZ07)]。
作者簡介: 王文采(1926-),研究员,中国科学院院士,著名植物分类学家,长期从事有花植物分类学研究。
Three new species of Delphinium L. (Ranunculaceae)
from Xinjiang
WANG Wen-tsai1, YANG Zongzong2
( 1. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy
of Sciences, Beijing 100093, China; 2. Yangs Herbarium, Urumqi 830011, China )
Abstract:Three species of the genus Dephinium L. (Ranunculaceae), D. buerjinense W. T. Wang & Z. Z. Yang and D. nilekeense W. T. Wang & Z. Z. Yang, belonging to Sect. Elatopsis Subsect. Elata, and D. habaheense W. T. Wang & Z. Z. Yang, belonging to Sect. Delphinastrum Subsect. Pogonantha, are described as new from Xinjiang Uygur Autonomous Region. The diagnostic differences between the three species and their allies are given respectively.
Key words: Ranunculaceae, Delphinium, new species, Xinjiang
布尔津翠雀花 图1
Delphinium buerjinense W. T. Wang & Z. Z. Yang, sp. nov. Fig. 1
(Sect.Elatopsis Huth Subsect. Elata W. T. Wang)
Ob racemos floribus 10-20 densissimis praeditos species nova haec est similis Delphinium gongliuensi W. T. Wang & Z. Z. Yang, a quo caulibus puberulis, foliorum laminis prope basin 3-partitis, racemis simplicibus, bracteolis anguste linearibus apice haud apiculatis, carpellis glabris praeclare differt. In D. gongliuensi, caules hirtelli, foliorum laminae leviter ultra medium 3-partitae, racemi compositi, bracteolae lineares apice longe apiculatae, et carpellorum ovaria puberula sunt.
Perennial herbs, 29-40 cm tall. Stems simple, puberulous, above base 4-5 mm across. Basal leaves 1-2; blades papery, pentagonal, 3 × 4.5-6 cm, at base deeply cordate, 3-parted to near base, central primary lobe obovate-rhombic, 3-lobed, with secondary lobes 1-2-lobulate, lateral primary lobes flabellate, unequally 3-parted, on both surfaces sparsely puberulous and mixed with glandular hairs; petioles 5-6 cm long. Cauline leaves similar to basal leaves, but larger, 4-6 × 9-10 cm, with central lobes rhombic or narrowly rhombic, and with petioles 2-7 cm long. Raceme terminal, 4-8 cm long, very densely 10-20-flowered; bracts linear, ca. 1 cm long; pedicels 0.3-1 cm long, with rachis and bracteoles densely puberulous and mixed with glandular hairs, above with narrow-linear bracteoles5-7 mm long. Flower: Sepals blue-purple, adaxially glabrous, abaxially puberulous and mixed with short glandular hairs, limb of upper sepal narrow-ovate,14 × 6 mm, spur subulate, 14-15 mm long, lateral sepals long elliptic, 15 × 6 mm, lower sepals obliquely obovate, 13×8 mm. Petals 2, black, glabrous. Staminodes 2, black; limbs oblong-elliptic, 6 × 3.5 mm, 2-lobed, on the middle densely barbate; claws 6 mm long. Stamens numerous, glabrous; filaments 6 mm long, below narrow-triangular, above filiform; anthers 1.5 × 0.8 mm. Carpels 3, glabrous; ovaries 2 × 0.8 mm, with inconspicuous netted nerves; styles1.1-1.5 mm long.
新疆(Xinjiang):布爾津县,喀纳斯湖(Buerjin County, Kanas Lake), alt. 1 820 m,山坡草地,萼片蓝紫色,有腺毛(in grassy place on slope, sepals blue-purple, with glandular hairs), 2020-07-06, 杨宗宗(Yang Zongzong) 3191 (holotype, PE; isotype, Yangs Herbarium);同地(seme locality), alt.1 820 m, 山坡草地,萼片近紫色 (in grassy place on slope, sepals subpurple), 2020-07-06,杨宗宗3190 (paratype, Yangs Herbarium)。
此种的总状花序有10~20朵极为密集的花,因此与也具此特征的巩留翠雀花(Delphinium gongliuense W. T. Wang & Z. Z. Yang)相似,与后者的区别在于此种的茎被短柔毛,叶片3深裂达近基部,总状花序简单,小苞片狭条形,顶端不具细尖头,心皮无毛。在巩留翠雀花,茎被微硬毛,叶片3深裂稍超过中部,总状花序复杂,小苞片条形,顶端具长细尖头,心皮子房被短柔毛(王文采和杨宗宗,2020)。
尼勒克翠雀花 图2
Delphinium nilekeense W. T. Wang & Z. Z. Yang, sp. nov. Fig. 2
(Sect.Elatopsis Huth Subsect. Elata W. T. Wang)
Ob foliorum laminarum forma, staminodii limbum nigrum subquadratum 4 mm longum apice 2-lobulatum species nova haec est affinis Delphinium iliensi Huth, a quo caulibus dense hispidis, racemis compositis, pedicellis dense puberulis, sepalis persistentibus, carpellorum ovariis dense puberulis, seminibus dense transversaliter anguste lamellatis distinguitur. In D. iliense, caules sparse hispidi, racemi simplices, pedicelli glabri, carpellorum ovaria subglabra, semina tantum ad angulos anguste alata sunt.
Perennial herbs, 90-108 cm tall. Stems simple, above base 6-10 mm across, densely spreading-hispid. Basal leaves 1-2, long petiolate; blades papery, pentagonal, ca. 9×15 cm, 3-parted to 2 cm above base, central primary lobe rhombic, 4.5 cm broad, secondary lobes with few triangular lobules and acute teeth, lateral primary lobes obliquely flabellate, ca. 9 cm broad, unequally 2-cleft; surfaces adaxially and abaxially appressed-puberulous and mixed with glandular hairs; petioles 15-24 cm long. Lower cauline leaves similar to basal ones, and upper cauline leaves smaller and shortly petiolate. Compound raceme terminal, 24-38 cm long, terminal raceme 14-20 cm long, 14-28-flowered, lateral racemes 3-4, 7-10 cm long, densely 6-10-flowered; lower bracts similar to upper cauline leaves,
upper bracts smaller, 3-sect, with entire narrow- lanceolate segments; pedicels 0.4-2.8 cm long, with rachis and bracteoles densely puberulous and mixed with yellow glandular hairs, above or near apex with linear bracteoles 4-6 mm long. Flower: Sepals blue-purple, persistent, adaxially glabrous, abaxially densely appressed-puberulous and mixed with glandular hairs, limb of upper sepal broad-ovate, ca. 11-12×8.5 mm, spur subulate, 11-12 mm long, lateral sepals orbicular-ovate, ca. 13×9 mm, lower sepals broad-elliptic, ca. 12×7.5 mm. Petals 2, black, glabrous. Staminodes 2; limbs black, subquadrate, ca. 4 × 3.5 mm, on apex 2-lobulate, on the middle densely barbate, on above margin long ciliate; claws whitish, ca. 5 mm long. Stamens numerous, glabrous; filaments 7 mm long, below narrow-triangular, above filiform; anthers ca. 1.2×1 mm. Carpels 3; ovaries ellipsoidal, 4 × 2 mm, densely puberulous; styles 2.5 mm long, glabrous. Follicles narrow-ovoid, ca. 11×4 mm, densely puberulous; persistent styles 3 mm long, glabrous. Seeds black, inconspicuously cuneate-tetrahedral, 2 mm long, densely trangsversely narrow-lamellate.
新疆 (Xinjiang):尼勒克縣,独库公路,乔尔玛 (Nileke County, Duku High-way, Qiaoerma), alt.2 420 m,山坡草地,萼片蓝紫色,宿存,有黄色腺毛 (in grassy place on slope, sepals blue-purple, persistent, with yellow glandular hairs), 2019-08-07,杨宗宗 (Yang Zongzong) 2531 (holotype, PE; isotype, Yangs Herbarium);同地 (seme locality), alt. 2 380 m,山坡,萼片蓝紫色,混生腺毛 (on slope, sepals blue-purple, mixed with glandular hairs),2019-08-07,杨宗宗 2538(paratype, Yangs Herbarium)。
此种的叶片五角形,3深裂,退化雄蕊瓣片黑色,近方形,顶端2微裂,因此与也具上述特征的伊犁翠雀花(Delphinium iliense Huth)在亲缘关系上相近;与后者的区别在于此种的茎密被糙硬毛,总状花序复杂,花梗密被短柔毛,萼片宿存,心皮子房也密被短柔毛,种子具横向密集的狭片。……